Shuning earned her PhD from Rensselaer Polytechnic Institute in 2007. Her thesis work was published as
Shuning Zhang, Gina M. Dailey, Elaine Kwan, Bernadette M. Glasheen, Gyna E. Sroga, and Andrea Page-McCaw. 2006. An MMP Liberates the Ninjurin A Ectodomain to Signal a Loss of Cell Adhesion. Genes & Development 20, 1899-1910.